:root {
  --ink: #17211d;
  --muted: #66736c;
  --line: #d8ddd7;
  --paper: #f7f6f0;
  --surface: #fff;
  --leaf: #2f6f4e;
  --water: #2d6f86;
  --coral: #c96f5b;
  --shade: rgba(23, 33, 29, .1);
}

* {
  box-sizing: border-box;
}

body {
  margin: 0;
  color: var(--ink);
  background: var(--paper);
  font-family: -apple-system, BlinkMacSystemFont, "Hiragino Sans", "Yu Gothic", "YuGothic", sans-serif;
  line-height: 1.7;
}

a {
  color: inherit;
  text-decoration-color: rgba(47, 111, 78, .35);
  text-underline-offset: .18em;
}

img {
  max-width: 100%;
  height: auto;
}

.skip-link {
  position: absolute;
  left: -999px;
}

.skip-link:focus {
  left: 1rem;
  top: 1rem;
  z-index: 99;
  padding: .6rem .8rem;
  background: var(--surface);
}

.site-header {
  position: sticky;
  top: 0;
  z-index: 20;
  border-bottom: 1px solid var(--line);
  background: rgba(247, 246, 240, .93);
  backdrop-filter: blur(14px);
}

.site-header__inner,
.site-footer__inner {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 1.5rem;
  max-width: 1180px;
  margin: 0 auto;
  padding: .9rem clamp(1rem, 4vw, 2.5rem);
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: .65rem;
  color: var(--ink);
  font-weight: 800;
  text-decoration: none;
}

.brand__mark {
  display: grid;
  place-items: center;
  width: 2rem;
  height: 2rem;
  border-radius: 999px;
  color: var(--paper);
  background: var(--ink);
}

.primary-nav ul,
.site-footer ul {
  display: flex;
  align-items: center;
  gap: 1rem;
  margin: 0;
  padding: 0;
  list-style: none;
}

.primary-nav a,
.site-footer a {
  color: var(--muted);
  font-size: .95rem;
  text-decoration: none;
}

.site-main {
  min-height: 72vh;
}

.hero {
  display: grid;
  grid-template-columns: minmax(0, .9fr) minmax(320px, 1.1fr);
  gap: clamp(1.5rem, 5vw, 4rem);
  align-items: center;
  max-width: 1180px;
  margin: 0 auto;
  padding: clamp(2rem, 7vw, 5rem) clamp(1rem, 4vw, 2.5rem) 2rem;
}

.hero h1,
.species-hero h1,
.entry-header h1,
.archive-header h1,
.not-found h1 {
  margin: 0 0 1rem;
  font-size: clamp(2.35rem, 5vw, 4.5rem);
  line-height: 1.08;
  letter-spacing: 0;
}

.hero p {
  max-width: 42rem;
  color: var(--muted);
  font-size: 1.08rem;
}

.hero img {
  display: block;
  width: 100%;
  aspect-ratio: 16 / 10;
  object-fit: cover;
  border-radius: 8px;
  box-shadow: 0 1.5rem 3rem var(--shade);
}

.eyebrow {
  margin: 0 0 .6rem;
  color: var(--coral);
  font-size: .82rem;
  font-weight: 800;
  letter-spacing: 0;
  text-transform: uppercase;
}

.hero-search {
  display: flex;
  gap: .5rem;
  max-width: 42rem;
  margin-top: 1.5rem;
}

.hero-search label {
  flex: 1;
}

.hero-search input[type="search"] {
  width: 100%;
  min-height: 3rem;
  padding: .7rem .9rem;
  border: 1px solid var(--line);
  border-radius: 6px;
  background: var(--surface);
  color: var(--ink);
  font: inherit;
}

.hero-search button,
.button-link {
  min-height: 3rem;
  padding: .7rem 1rem;
  border: 0;
  border-radius: 6px;
  background: var(--leaf);
  color: #fff;
  font: inherit;
  font-weight: 700;
  text-decoration: none;
  cursor: pointer;
}

.section-band,
.archive-header,
.post-list,
.content-shell,
.note-principles {
  max-width: 1180px;
  margin: 0 auto;
  padding: clamp(2rem, 5vw, 4rem) clamp(1rem, 4vw, 2.5rem);
}

.section-heading {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 1rem;
  margin-bottom: 1.25rem;
}

.section-heading h2 {
  margin: 0;
  font-size: clamp(1.8rem, 3vw, 2.6rem);
}

.species-card-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1rem;
}

.species-card,
.post-list__item {
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: 8px;
  background: var(--surface);
}

.species-card__thumb img {
  display: block;
  width: 100%;
  aspect-ratio: 4 / 3;
  object-fit: cover;
}

.species-card__body,
.post-list__item {
  padding: 1.25rem;
}

.species-card__meta,
.post-meta,
.species-summary {
  color: var(--muted);
  font-size: .92rem;
}

.species-card h3,
.post-list h2 {
  margin: 0 0 .45rem;
  font-size: 1.35rem;
  line-height: 1.35;
}

.species-card p,
.post-list p,
.empty-state {
  margin: 0;
  color: var(--muted);
}

.note-principles {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1px;
}

.note-principles article {
  min-height: 12rem;
  padding: 1.4rem;
  border: 1px solid var(--line);
  background: #eef2ea;
}

.note-principles span {
  color: var(--coral);
  font-weight: 900;
}

.note-principles h2 {
  margin: .6rem 0;
  font-size: 1.3rem;
}

.species-hero {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(260px, 420px);
  gap: 2rem;
  align-items: end;
  max-width: 1180px;
  margin: 0 auto;
  padding: clamp(2rem, 6vw, 4.5rem) clamp(1rem, 4vw, 2.5rem) 1.5rem;
}

.species-hero__image img {
  display: block;
  width: 100%;
  aspect-ratio: 4 / 3;
  object-fit: cover;
  border-radius: 8px;
}

.entry-content {
  max-width: 850px;
  font-size: 1.04rem;
}

.entry-content h2,
.entry-content h3 {
  margin-top: 2rem;
  line-height: 1.3;
}

.site-footer {
  border-top: 1px solid var(--line);
  color: var(--muted);
}

.site-footer__inner {
  align-items: flex-start;
}

@media (max-width: 900px) {
  .hero,
  .species-hero {
    grid-template-columns: 1fr;
  }

  .species-card-grid,
  .note-principles {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }
}

@media (max-width: 640px) {
  .site-header__inner,
  .site-footer__inner,
  .hero-search {
    align-items: stretch;
    flex-direction: column;
  }

  .species-card-grid,
  .note-principles {
    grid-template-columns: 1fr;
  }
}